Ichthyovenator


Ichthyovenator is a genus of spinosaurid dinosaur that lived in what is now Laos, sometime between 125 and 113 million years ago, during the Aptian stage of the Early Cretaceous period. It is known from fossils collected from the Grès supérieurs Formation of the Savannakhet Basin, the first of which were found in 2010, consisting of a partial skeleton without the skull or limbs. This specimen became the holotype of the new genus and species Ichthyovenator laosensis, and was described by palaeontologist Ronan Allain and colleagues in 2012. The generic name, meaning "fish hunter", refers to its assumed piscivorous lifestyle, while the specific name alludes to the country of Laos. In 2014, it was announced that more remains from the dig site had been recovered; these fossils included teeth, more vertebrae (backbones) and a pubic bone from the same individual.

The holotype specimen is estimated to have been between 8.5 to 10.5 metres (28 to 34 feet) long and to have weighed 2.4 tonnes (2.6 short tons; 2.4 long tons). Ichthyovenator's teeth were straight and conical, and its neck resembled that of the closely related genus Sigilmassasaurus. Like others in its family, Ichthyovenator had tall neural spines that formed a sail on its back. Unlike other known spinosaurids, Ichthyovenator's sail had a sinusoidal (wave-like) shape that curved downwards over the hips and divided into two separate sails. The pelvic girdle was reduced; the ilium—the uppermost body of the pelvis—was proportionately longer than both the pubis and ischium than in other known theropod dinosaurs. Ichthyovenator was initially thought to belong to the subfamily Baryonychinae but more recent analyses place it as a primitive member of the Spinosaurinae.

As a spinosaur, Ichthyovenator would have had a long, shallow snout and robust forelimbs. Its diet likely mainly consisted of aquatic prey, hence its etymology. Spinosaurids are also known to have eaten small dinosaurs and pterosaurs in addition to fish. Ichthyovenator's conspicuous sail might have been used for sexual display or species recognition. Fossil evidence suggests spinosaurids, especially spinosaurines, were adapted for semiaquatic lifestyles. The vertebral spines of Ichthyovenator's tail were unusually tall, suggesting—as in today's crocodilians—the tail may have aided in swimming. Ichthyovenator lived alongside sauropod and ornithopod dinosaurs, as well as bivalves, fish and turtles.

The first fossils of Ichthyovenator were found in 2010 at Ban Kalum in the Grès supérieurs Formation of the Savannakhet Basin in Savannakhet Province, Laos. These fossilized bones were recovered from a red sandstone layer within a surface area of less than 2 square metres (22 sq ft). Designated under the specimen numbers MDS BK10-01 to 15, they consist of a partly articulated, well-preserved skeleton lacking the skull and limbs, and including the third-to-last dorsal (back) vertebra, the neural spine of the last dorsal vertebra, five partial sacral (hip) vertebrae, the first two caudal (tail) vertebrae, both ilia (main hip bones), a right pubis (pubic bone), both ischia (lower and rearmost hip bones) and a posterior dorsal rib. The twelfth dorsal spine is bent sideways when viewed front-to-back due to taphonomic distorsion. The centra (vertebral bodies) of the sacrals are largely incomplete due to erosion, but preserved all of their accompanying spines with their upper edges intact. At the time of Ichthyovenator's description, excavations at the site were still ongoing.[1]


Skeletal diagram with holotype fossils in white and undescribed material in red
Casts of the vertebrae at the Muséum National d'Histoire Naturelle, Paris
Size of various spinosaurids (Ichthyovenator in turquoise, first from right) compared with a human
Casts of the known spinal column at the National Museum of Nature and Science, Tokyo. The thirteenth dorsal spine and the lower part of the sacrum have been partially reconstructed.
Various caudal (tail) vertebrae and chevrons from the holotype
Speculative life restoration, with missing elements such as the head and limbs based on relatives
Comparison of the pelvic region and neural spine sails of Suchomimus, Spinosaurus, and Ichthyovenator
Reconstructed skeleton of the spinosaurine Irritator, mounted at the National Museum of Nature and Science, Tokyo
Casts of the vertebrae seen from behind
Ichthyovenator might have used its tail, which had elongated neural spines, for propulsion through water, similar to crocodilians such as this Nile crocodile[15]
Fossils of the sauropod Tangvayosaurus, another dinosaur from the Gres superieurs Formation, Savannakhet Dinosaur Museum