Praemegaceros is an extinct genus of deer, known from the Pleistocene and Holocene of Western Eurasia. It contains the subgenera Praemegaceros, Orthogonoceros and Nesoleipoceros. It has sometimes been synonymised with Megaloceros and Megaceroides, however they have been found to be generically distinct.[1]
P. obscurus is the earliest known species from the Early Pleistocene of Europe, and had long, crooked antlers. P. verticornis is an Early to Mid-Pleistocene species, closely related to P. obscurus, which lived throughout Southern Europe.[1][2] The genus was widely distributed across Europe, West and Central Asia during the Early-Middle Pleistocene, with fossils having been discovered in France, Georgia, Germany, England, Greece, Israel, Italy, Romania, Russia Spain, Syria, and Tajikistan.[3][4][5] The genus was extinct in mainland Europe and Asia by end of the Middle Pleistocene. An insular species, P. cazioti survived into the Late Pleistocene and Holocene in isolation on Sardinia until around 5500 BCE.[6]
Praemegaceros was first described as a subgenus of Cervus by Portis in 1920, to include Cervus (Praemegaceros) dawkinsi. Kalkhe in 1956 named Orthogonoceros with the type species of Orthogonoceros (previously Cervus) verticornis. Kalkhe in 1965 recognised that these genera were synonyms, with Praemegaceros having priority. Radulesco & Samson in 1967 designated P. dawkinsi as the type species of the genus, while also naming the new genera Allocaenelaphus, Psecupsoceros, and Nesoleipoceros which are now recognised as synonyms of the genus.[7]
Known from the late Villafranchian of Europe (including Central Italy, Central Romania and Moldova) and the Near East, extending from the Cromer Forest Bed in England to Ubeidiya, Israel, Dmanisi, Georgia and the Azov region, Russia. Croitor suggests an origin in South Asia for this species, descended from taxa possibly referrable to Panolia sp. from the Siwaliks.[7]
Named in 1882 from remains from the Middle Pleistocene Cromer Forest Bed in Norfolk by Edwin Tulley Newton which were originally described in 1872 as belonging to P. verticornis by William Boyd Dawkins. The size is estimated to be around 220 kg, with a mesodont dentition. Croitor suggests that because the pedicles (base of the antlers) are robust and similar to those of giant deer, that P. dawkinsi represents a dwarfed form.[7]
Named by Wolfgang Soergel in 1927 for the species found in the lowest level of the Middle Pleistocene Mosbach locality in Germany.[8] It was included in the “verticornis” group of Azzaroli's 1953 classification. It has been suggested to a be synonym of P. verticornis. However, Croitor suggests that these similarities are the result of parallel evolution, and proposes that P. mosbachensis represents an intermediate form between P. obscurus and P. dawkinsi.[7]