Noctuidae


The Noctuidae, commonly known as owlet moths, cutworms or armyworms, are a family of moths. They are considered the most controversial family in the superfamily Noctuoidea because many of the clades are constantly changing, along with the other families of the Noctuoidea.[1][2][3] It was considered the largest family in Lepidoptera for a long time, but after regrouping Lymantriinae, Catocalinae and Calpinae within the family Erebidae, the latter holds this title now.[4] Currently, Noctuidae is the second largest family in Noctuoidea, with about 1,089 genera and 11,772 species.[5] This classification is still contingent, as more changes continue to appear between Noctuidae and Erebidae.

Adult: Most noctuid adults have drab wings, but some subfamilies, such as Acronictinae and Agaristinae, are very colorful, especially those from tropical regions (e.g. Baorisa hieroglyphica). They are characterized by a structure in the metathorax called the nodular sclerite or epaulette, which separates the tympanum and the conjunctiva in the tympanal organ. It functions to keep parasites (Acari) out of the tympanal cavity. Another characteristic in this group is trifine hindwing venation, by reduction or absence of the second medial vein (M2).[6]

Markings present on the wings of noctuid adults can be helpful in distinguishing species. From the basal location to the outer edge (proximal to distal) on the forewing, there is a claviform (club-shaped) stigma, horizontally oriented with the thicker end closer to the wing's outer edge, located posterior to a discal (round) stigma.[7] These are followed distally by a reniform (kidney-shaped) stigma,[8] which is typically oriented with its concave side facing the wing's outer edge. It is often not possible to discern all of the stigmata on all specimens or species.[7] Crossbands or crosslines may be present, oriented longitudinally from the leading to the trailing edge of the wing.[8]

Larva: Commonly green or brown; some species present bright colors, such as the camphorweed cucullia moth (Cucullia alfarata). Most are pudgy and smooth with rounded short heads and few setae, but there are some exceptions in some subfamilies (e.g. Acronictinae and Pantheinae).[9]

Pupa: The pupae most often range from shiny brown to dark brown. When they newly pupate they are bright brownish orange, but after a few days start to get darker.

The word Noctuidae is derived from the name of the type genus Noctua, which is the Latin name for the little owl, and the patronymic suffix -idae used typically to form taxonomic family names in animals.[10]


Noctuidae wings venation
Noctuidae wings venation
Setaceous Hebrew character
A Lychnis moth caterpillar feeding on the seeds of red campion (Silene dioica).
The eight-spotted forester moth (Alypia octomaculata) puddling on water from a leaf of firebush (Croton lucidus).
Lesser yellow underwing
The Spanish moth feeding on Amaryllis sp.
The Old World bollworm caterpillar feeding on a strawberry.