Siamosaurus


Siamosaurus (meaning "Siam reptile") is a genus of spinosaurid dinosaur that lived in what is now known as China and Thailand during the Early Cretaceous period (Barremian to Aptian) and is the first reported spinosaurid from Asia. It is confidently known only from tooth fossils; the first were found in the Sao Khua Formation, with more teeth later recovered from the younger Khok Kruat Formation. The type species Siamosaurus suteethorni, whose name honours Thai palaeontologist Varavudh Suteethorn, was formally described in 1986. In 2009, four teeth from China previously attributed to a pliosaur—under the species "Sinopliosaurus" fusuiensis—were identified as those of a spinosaurid, possibly Siamosaurus. It is yet to be determined if two partial spinosaurid skeletons from Thailand and an isolated tooth from Japan also belong to Siamosaurus.

Since it is based only on teeth, Siamosaurus's body size is uncertain, though it has been estimated at between 5.1 to 9.1 metres (17 to 30 feet) in length. The holotype tooth is 62.5 millimetres (2.46 inches) long. Siamosaurus's teeth were straight, oval to circular in cross-section, and lined with distinct lengthwise grooves. Its teeth had wrinkled enamel, similar to teeth from the related genus Baryonyx. As a spinosaur it would have had a long, low snout and robust forelimbs, and one possible skeleton indicates the presence of a tall sail running down its back, another typical trait of this theropod family. Siamosaurus is considered by some palaeontologists to be a dubious name, with some arguing that its teeth are hard to differentiate from those of other Early Cretaceous spinosaurids, and others that it may not be a dinosaur at all. Based on dental traits, Siamosaurus and "S." fusuiensis have been placed in the subfamily Spinosaurinae.

Like in all spinosaurids, Siamosaurus's teeth were conical, with reduced or absent serrations. This made them suitable for impaling rather than tearing flesh, a trait typically seen in largely piscivorous (fish-eating) animals. Spinosaurids are also known to have consumed pterosaurs and small dinosaurs, and there is fossil evidence of Siamosaurus itself feeding on sauropod dinosaurs, either via scavenging or active hunting. Siamosaurus's role as a partially piscivorous predator may have reduced the prominence of some contemporaneous crocodilians competing for the same food sources. Isotope analysis of the teeth of Siamosaurus and other spinosaurids indicates semiaquatic habits. Siamosaurus lived in a semi-arid habitat of floodplains and meandering rivers, where it coexisted with other dinosaurs, as well as pterosaurs, fishes, turtles, crocodyliforms, and other aquatic animals.

The Sao Khua Formation, where the first Siamosaurus fossils were discovered, is part of the Khorat Group.[1] The formation is dated to the Barremian stage of the Early Cretaceous period, 129.4 to 125 million years ago.[2][3] In 1983, French palaeontologist Éric Buffetaut and his Thai colleague Rucha Ingavat described a set of fossil teeth recovered from the Phu Pratu Teema locality of the Sao Khua Formation, in the Phu Wiang area of Khon Kaen Province. They did not conclude as to what animal they originated from, their opinion being that the specimens belonged "either to an unusual theropod dinosaur or to some unknown crocodilian". In 1986, a reassessment of the remains by the same authors attributed them to a new genus and species of spinosaurid theropod, which they named Siamosaurus suteethorni. The generic name alludes to the ancient name of Thailand, "Siam", and is combined with the Ancient Greek word σαῦρος (sauros), meaning "lizard" or "reptile". The specific name honours Thai geologist and palaeontologist Varavudh Suteethorn, and his contributions to vertebrate palaeontology discoveries in Thailand.[1]


Map of northeastern Thailand, with outcrops of the Sao Khua Formation (dark gray) and the Khok Kruat and Phu Kradung Formations (light gray)
Sculptures of Siamosaurus at the Sirindhorn Museum (top) and Phu Wiang Dinosaur Museum (bottom); both buildings also house fossils of the animal.[10][11][13]
Illustration of caudal (tail) vertebrae of "Phuwiang spinosaurid B" from the Sao Khua Formation, whose remains are potentially attributable to S. suteethorni
Tentatively estimated size of Siamosaurus (in gray) and "Phuwiang spinosaurid B" (in red), which may represent the same taxon
Diagram comparing the holotype tooth (first from left) with other spinosaurid teeth from Asia
Partial tooth of an indeterminate spinosaurid (KDC-PV-0003) and possible Siamosaurus tooth (GMNH-PV-999), in the National Museum of Nature and Science, Tokyo
Skeletal diagram showing known elements of spinosaurid specimen SM-KK 14
Tooth of the related genus Spinosaurus, Museo di Storia Naturale A. Stoppani, Lombardy
Spinosaurid teeth are often mistaken for those of plesiosaurs (above) and vice versa, though there are certain differences between their dentition.
Restoration of Ichthyovenator, a spinosaurine from what is now Laos and one of the closest known relatives of Siamosaurus.[38]
Vertebra from specimen SM-KK14, which may belong to Siamosaurus
Speculative life restoration of Siamosaurus (right) in the Sao Khua Formation environment, with the goniopholidid Sunosuchus (middle left) and a herd of the sauropods Phuwiangosaurus in the background
Annotated skull diagram of the related Spinosaurus
Restoration of a generic spinosaur feeding on a sauropod carcass in the Sao Khua Formation environment, a behavior hypothesized based on teeth belonging to Siamosaurus. The ornithomimosaurs in the background are Kinnareemimus.[5][63]
Geological map of the Khorat Plateau in northeast Thailand