Allosaurus


Allosaurus (/ˌæləˈsɔːrəs/[2][3]) is an extinct genus of large carnosaurian theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic epoch (Kimmeridgian to late Tithonian). The name "Allosaurus" means "different lizard" alluding to its unique (at the time of its discovery) concave vertebrae. It is derived from the Greek ἄλλος (allos) ("different, other") and σαῦρος (sauros) ("lizard / generic reptile"). The first fossil remains that could definitively be ascribed to this genus were described in 1877 by paleontologist Othniel Charles Marsh. As one of the first well-known theropod dinosaurs, it has long attracted attention outside of paleontological circles.

Allosaurus was a large bipedal predator. Its skull was light, robust and equipped with dozens of sharp, serrated teeth. It averaged 9.5 meters (31 ft) in length for A. fragilis, with the maximum length estimate being 11.5 meters long. Relative to the large and powerful hindlimbs, its three-fingered forelimbs were small, and the body was balanced by a long and heavily muscled tail. It is classified as an allosaurid, a type of carnosaurian theropod dinosaur.

The genus has a complicated taxonomy, and includes three valid species, the best known of which is A. fragilis. The bulk of Allosaurus remains have come from North America's Morrison Formation, with material also known from Portugal. It was known for over half of the 20th century as Antrodemus, but a study of the copious remains from the Cleveland-Lloyd Dinosaur Quarry brought the name "Allosaurus" back to prominence and established it as one of the best-known dinosaurs.

As the most abundant large predator in the Morrison Formation, Allosaurus was at the top of the food chain, probably preying on contemporaneous large herbivorous dinosaurs, and perhaps other predators. Potential prey included ornithopods, stegosaurids, and sauropods. Some paleontologists interpret Allosaurus as having had cooperative social behavior, and hunting in packs, while others believe individuals may have been aggressive toward each other, and that congregations of this genus are the result of lone individuals feeding on the same carcasses.

Allosaurus was a typical large theropod, having a massive skull on a short neck, a long, slightly sloping tail, and reduced forelimbs. Allosaurus fragilis, the best-known species, had an average length of 8.5 m (28 ft),[4] with the largest definitive Allosaurus specimen (AMNH 680) estimated at 9.7 meters (32 feet) long,[5] with an estimated weight of 2.3 metric tons (2.5 short tons).[5] In his 1976 monograph on Allosaurus, James H. Madsen mentioned a range of bone sizes which he interpreted to show a maximum length of 12 to 13 m (39 to 43 ft).[6] As with dinosaurs in general, weight estimates are debatable, and since 1980 have ranged between 1,500 kilograms (3,300 pounds), 1,000 to 4,000 kg (2,200 to 8,800 lb), and 1,010 kilograms (2,230 pounds) for modal adult weight (not maximum).[7] John Foster, a specialist on the Morrison Formation, suggests that 1,000 kg (2,200 lb) is reasonable for large adults of A. fragilis, but that 700 kg (1,500 lb) is a closer estimate for individuals represented by the average-sized thigh bones he has measured.[8] Using the subadult specimen nicknamed "Big Al", since assigned to the species Allosaurus jimmadseni,[1] researchers using computer modelling arrived at a best estimate of 1,500 kilograms (3,300 lb) for the individual, but by varying parameters they found a range from approximately 1,400 kilograms (3,100 lb) to approximately 2,000 kilograms (4,400 lb).[9]


The size range of Allosaurus compared with a human
A. jimmadseni skeletal reconstruction
A. jimmadseni skull with diagram highlighting individual bones
Life restoration of A. fragilis
Hand and claws of A. fragilis
AMNH 5753 in a Charles R. Knight life restoration
A. fragilis at the Cleveland-Lloyd Dinosaur Quarry museum, Utah
"Big Al" at the Museum of the Rockies
Diagram comparing skulls of the three recognized species; A. fragilis (A), A. jimmadseni (B), A. europeaus (C)
Holotype material of Creosaurus atrox
Antrodemus valens holotype tail vertebra (above) compared to the same of Allosaurus (below)
A. tendagurensis tibia, Naturkunde Museum Berlin
Restored skeleton of Saurophaganax or A. maximus
Mounted skeleton of "Big Al II" (specimen SMA 0005)
Skeletons at different growth stages on display, the Natural History Museum of Utah
Restoration of a juvenile Allosaurus
Allosaurus and Stegosaurus skeletons, the Denver Museum of Nature and Science
Bitten Stegosaurus plate close-up, showing how well the damage matches the front of an Allosaurus "mouth"
A. fragilis showing its maximum possible gape, based on Bakker (1998) and Rayfield et al. (2001)
Illustrations showing the skull of A. jimmadseni from the side (A), top (B), and back (C)
The holotype dentary of Labrosaurus ferox, which may have been injured by the bite of another A. fragilis
Endocast (cast of the brain cavity) of Allosaurus
Mounted A. fragilis skeleton (USNM4734), which has several healed injuries
Skeletal restoration of "Big Al II" showing bones with pathologies
Restoration of Barosaurus rearing to defend itself against a pair of A. fragilis
Locations in the Morrison Formation (yellow) where Allosaurus remains have been found
Dry season at the Mygatt-Moore Quarry showing Ceratosaurus (center) and Allosaurus fighting over the desiccated carcass of another theropod