Archaeopteryx


Archaeopteryx (/ˌɑːrkˈɒptərɪks/; lit.'old-wing'), sometimes referred to by its German name, "Urvogel" (lit. 'original bird' or 'first bird'), is a genus of bird-like dinosaurs. The name derives from the ancient Greek ἀρχαῖος (archaīos), meaning "ancient", and πτέρυξ (ptéryx), meaning "feather" or "wing".Between the late 19th century and the early 21st century, Archaeopteryx was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae).[2] Older potential avialans have since been identified, including Anchiornis, Xiaotingia, and Aurornis.[3]

Archaeopteryx lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven,[4] the largest species of Archaeopteryx could grow to about 0.5 m (1 ft 8 in) in length. Despite their small size, broad wings, and inferred ability to fly or glide, Archaeopteryx had more in common with other small Mesozoic dinosaurs than with modern birds. In particular, they shared the following features with the dromaeosaurids and troodontids: jaws with sharp teeth, three fingers with claws, a long bony tail, hyperextensible second toes ("killing claw"), feathers (which also suggest warm-bloodedness), and various features of the skeleton.[5][6]

These features make Archaeopteryx a clear candidate for a transitional fossil between non-avian dinosaurs and birds.[7][8] Thus, Archaeopteryx plays an important role, not only in the study of the origin of birds, but in the study of dinosaurs. It was named from a single feather in 1861,[9] the identity of which has been controversial.[10][11] That same year, the first complete specimen of Archaeopteryx was announced. Over the years, ten more fossils of Archaeopteryxhave surfaced. Despite variation among these fossils, most experts regard all the remains that have been discovered as belonging to a single species, although this is still debated.

Archaeopteryx was long considered to be the beginning of the evolutionary tree of birds. It has qualities that helped define what it is like to be a bird, such as its long, powerful front limbs. However, in recent years, the discovery of several small, feathered dinosaurs has created a mystery for paleontologists, raising questions about which animals are the ancestors of modern birds and which are their relatives.[12]

Most of these eleven fossils include impressions of feathers. Because these feathers are of an advanced form (flight feathers), these fossils are evidence that the evolution of feathers began before the Late Jurassic.[13] The type specimen of Archaeopteryx was discovered just two years after Charles Darwin published On the Origin of Species. Archaeopteryx seemed to confirm Darwin's theories and has since become a key piece of evidence for the origin of birds, the transitional fossils debate, and confirmation of evolution.

Over the years, twelve body fossil specimens of Archaeopteryx have been found. All of the fossils come from the limestone deposits, quarried for centuries, near Solnhofen, Germany.[14][15]


Timeline of Archaeopteryx discoveries until 2007
The single feather
Cast of the Maxberg Specimen
Eichstätt Specimen, once considered a distinct genus, Jurapteryx
Daiting Specimen, the holotype of A. albersdoerferi
Bürgermeister-Müller ("chicken wing") Specimen
Details of the Wyoming Dinosaur Center Archaeopteryx (WDC-CSG-100)
The eleventh specimen
Specimens compared to a human in scale
Anatomical illustration comparing the "frond-tail" of Archaeopteryx with the "fan-tail" of a modern bird
Artist's restoration illustrating one interpretation of Carney's study[64]
The Thermopolis Specimen
The twelfth specimen
Skeletal restorations of various specimens
The Solnhofen Specimen, by some considered as belonging to the genus Wellnhoferia
Comparison of the forelimb of Archaeopteryx (right) with that of Deinonychus (left)
1880 photo of the Berlin specimen, showing leg feathers that were removed subsequently, during preparation
The Munich Specimen
Replica of the London Specimen
Reconstructed skeleton, Academy of Natural Sciences of Drexel University
Growth trends compared with other dinosaurs and birds
Restoration of Archaeopteryx chasing a juvenile Compsognathus