Panoplosaurus


Panoplosaurus is a genus of armoured dinosaur from the Late Cretaceous of Alberta, Canada. Few specimens of the genus are known, all from the middle Campanian of the Dinosaur Park Formation, roughly 76 to 75 million years ago. It was first discovered in 1917, and named in 1919 by Lawrence Lambe, named for its extensive armour, meaning "well-armoured lizard". Panoplosaurus has at times been considered the proper name for material otherwise referred to as Edmontonia, complicating its phylogenetic and ecological interpretations, at one point being considered to have existed across Alberta, New Mexico and Texas, with specimens in institutions from Canada and the United States. The skull and skeleton of Panoplosaurus are similar to its relatives, but have a few significant differences, such as the lumpy form of the skull osteoderms, a completely fused shoulder blade, and regularly shaped plates on its neck and body lacking prominent spines. It was a quadrupedal animal, roughly 5 m (16 ft) long and 1,600 kg (3,500 lb) in weight. The skull has a short snout, with a very domed surface, and bony plates directly covering the cheek. The neck had circular groups of plates arranged around the top surface, both the forelimb and hindlimb were about the same length, and the hand may have only included three fingers. Almost the entire surface of the body was covered in plates, osteoderms and scutes of varying sizes, ranging from large elements along the skull and neck, to smaller, round bones underneath the chin and body, to small ossicles that filled in the spaces between other, larger osteoderms.

Panoplosaurus was originally classified as a stegosaur related to the similarly-armoured form Ankylosaurus, a group that was later divided with ankylosaurs becoming their own group. It was then considered close to Edmontonia in the subfamily Panoplosaurinae, but then moved into a general placement in Nodosauridae. Edmontonia was for a time considered the same taxon as Panoplosaurus, making it the only nodosaur from the Campanian of North America, but this was quickly disputed and they are now considered separate. Following consistent placements in phylogenetic analyses close to Edmontonia and the American taxon Animantarx, Panoplosaurus was placed into the clade Panoplosaurini, related but not close to Nodosaurus or Struthiosaurus, which it was considered close to around when it was named. Panoplosaurus is from deposits slightly younger than Edmontonia rugosidens, and existed alongside hadrosaurids like Corythosaurus and Lambeosaurus, ceratopsids like Centrosaurus, and the tyrannosaurid Gorgosaurus, as well as other small dinosaurs like Stegoceras, Dromaeosaurus and Ornithomimus, and various fishes, amphibians, crocodiles and pterosaurs.


Outcrops of the Dinosaur Park Formation along the Red Deer River
Life restoration
Skull of Panoplosaurus mirus CMN 2759 in dorsal, ventral, and anterior views
Scapulocoracoid, humerus, and tibia and fibula of Panoplosaurus holotype CMN 2759
Osteoderms and arrangement of neck osteoderms of Panoplosaurus mirus
Skull of Edmontonia rugosidens in the RTMP, at times considered a species of Panoplosaurus
Mounted skeletons of Denversaurus and Tyrannosaurus, Houston Museum of Natural Science
Skulls of Panoplosaurus (left) and Euoplocephalus (right) and their respective sinuses
Map of North America and the Western Interior Seaway 75 mya
Depiction of megaherbivores in the Dinosaur Park Formation, Panoplosaurus on the far right