Salamander


Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten extant salamander families are grouped together under the order Urodela. Salamander diversity is highest in the Northern Hemisphere and most species are found in the Holarctic realm, with some species present in the Neotropical realm.

Salamanders rarely have more than four toes on their front legs and five on their rear legs, but some species have fewer digits and others lack hind limbs. Their permeable skin usually makes them reliant on habitats in or near water or other cool, damp places. Some salamander species are fully aquatic throughout their lives, some take to the water intermittently, and others are entirely terrestrial as adults. They are capable of regenerating lost limbs as well as other damaged parts of their bodies. Researchers hope to reverse engineer the remarkable regenerative processes for potential human medical applications, such as brain and spinal cord injury treatment or preventing harmful scarring during heart surgery recovery.[2]

Members of the family Salamandridae are mostly known as newts and lack the costal grooves along the sides of their bodies typical of other groups. The skin of some species contains the powerful poison tetrodotoxin; these salamanders tend to be slow-moving and have bright warning coloration to advertise their toxicity. Salamanders typically lay eggs in water and have aquatic larvae, but great variation occurs in their lifecycles. Some species in harsh environments reproduce while still in the larval state.

The skin lacks scales and is moist and smooth to the touch, except in newts of the Salamandridae, which may have velvety or warty skin, wet to the touch. The skin may be drab or brightly colored, exhibiting various patterns of stripes, bars, spots, blotches, or dots. Male newts become dramatically colored during the breeding season. Cave species dwelling in darkness lack pigmentation and have a translucent pink or pearlescent appearance.[3]

Salamanders range in size from the minute salamanders, with a total length of 27 mm (1+18 in), including the tail, to the Chinese giant salamander which reaches 1.8 m (6 ft) and weighs up to 65 kg (145 lb). Most, however, are between 10 and 20 cm (4 and 8 in) in length.[4]

An adult salamander generally resembles a small lizard, having a basal tetrapod body form with a cylindrical trunk, four limbs, and a long tail. Except in the family Salamandridae, the head, body, and tail have a number of vertical depressions in the surface which run from the mid-dorsal region to the ventral area and are known as costal grooves. Their function seems to be to help keep the skin moist by channeling water over the surface of the body.[5]


X-ray image of salamander
Sirens have an eel-like appearance.
The front part of the olm's head carries sensitive chemo-, mechano-, and electroreceptors.
Biofluorescence can be observed across various salamander species
Salamanders need moist environments to respire through their skin.
A dissected view of the levatores arcuum muscles in a Necturus maculosus specimen. These (shown in the purple circles) move the external gills, as a means of respiration.
The head of a tiger salamander
A fire salamander's striking black and yellow pattern warns off predators
Sierra newt amplexus found in stream at Woolman Semester in Nevada County, California
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Embryonic development of a salamander, filmed in the 1920s
Neotenic axolotl, showing external gills
The threatened hellbender
Karaurus sharovi
A salamander unharmed in the fire, 1350
Ukiyo-e print by Utagawa Kuniyoshi (1797–1861) depicting a giant salamander being stabbed by the samurai Hanagami Danjō no jō Arakage