Copepod


Copepods (/ˈkpɪpɒd/; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat. Some species are planktonic (inhabiting sea waters), some are benthic (living on the ocean floor), a number of species have parasitic phases, and some continental species may live in limnoterrestrial habitats and other wet terrestrial places, such as swamps, under leaf fall in wet forests, bogs, springs, ephemeral ponds, and puddles, damp moss, or water-filled recesses (phytotelmata) of plants such as bromeliads and pitcher plants. Many live underground in marine and freshwater caves, sinkholes, or stream beds. Copepods are sometimes used as biodiversity indicators.

As with other crustaceans, copepods have a larval form. For copepods, the egg hatches into a nauplius form, with a head and a tail but no true thorax or abdomen. The larva molts several times until it resembles the adult and then, after more molts, achieves adult development. The nauplius form is so different from the adult form that it was once thought to be a separate species. The metamorphosis had, until 1832, led to copepods being misidentified as zoophytes or insects (albeit aquatic ones), or, for parasitic copepods, 'fish lice'.[1]

Copepods form a subclass belonging to class Hexanauplia in the subphylum Crustacea (crustaceans); they are divided into 10 orders. Some 13,000 species of copepods are known, and 2,800 of them live in fresh water.[2][3]

Copepods vary considerably, but are typically 1 to 2 mm (132 to 332 in) long, with a teardrop-shaped body and large antennae. Like other crustaceans, they have an armoured exoskeleton, but they are so small that in most species, this thin armour and the entire body is almost totally transparent. Some polar copepods reach 1 cm (12 in). Most copepods have a single median compound eye, usually bright red and in the centre of the transparent head; subterranean species may be eyeless. Like other crustaceans, copepods possess two pairs of antennae; the first pair is often long and conspicuous.

Free-living copepods of the orders Calanoida, Cyclopoida, and Harpacticoida typically have a short, cylindrical body, with a rounded or beaked head, although considerable variation exists in this pattern. The head is fused with the first one or two thoracic segments, while the remainder of the thorax has three to five segments, each with limbs. The first pair of thoracic appendages is modified to form maxillipeds, which assist in feeding. The abdomen is typically narrower than the thorax, and contains five segments without any appendages, except for some tail-like "rami" at the tip.[7] Parasitic copepods (the other seven orders) vary widely in morphology and no generalizations are possible.

Because of their small size, copepods have no need of any heart or circulatory system (the members of the order Calanoida have a heart, but no blood vessels), and most also lack gills. Instead, they absorb oxygen directly into their bodies. Their excretory system consists of maxillary glands.


Most copepods have a single naupliar eye in the middle of their head, but copepods of the genera Copilia and Corycaeus possess two eyes. Each eye has a large anterior cuticular lens paired with a posterior internal lens to form a telescope.[4][5][6]
Slow-motion macrophotography video (50%), taken using ecoSCOPE, of juvenile Atlantic herring (38 mm) feeding on copepods – the fish approach from below and catch each copepod individually. In the middle of the image, a copepod escapes successfully to the left.
Egg sac of a copepod
Lernaeolophus sultanus (Pennellidae), parasite of the fish Pristipomoides filamentosus, scale: each division = 1 mm [16]
Acanthochondria cornuta, an ectoparasite on flounder in the North Sea