Triceratops


Triceratops is an extinct genus of herbivorous chasmosaurine ceratopsid dinosaur that first appeared during the late Maastrichtian stage of the Late Cretaceous period, about 68 million years ago in what is now North America. It is one of the last-known non-avian dinosaur genera, and became extinct in the Cretaceous–Paleogene extinction event 66 million years ago. The name Triceratops, which literally means 'three-horned face', is derived from the Greek words trí- (τρί-) meaning 'three', kéras (κέρας) meaning 'horn', and ṓps (ὤψ) meaning 'face'.

Bearing a large bony frill, three horns on the skull, and a large four-legged body, exhibiting convergent evolution with rhinoceroses and bovines, Triceratops is one of the most recognizable of all dinosaurs and the most well-known ceratopsid. It was also one of the largest, up to 9 meters (29.5 ft) long and 12 metric tons (13 short tons) in weight. It shared the landscape with and was most likely preyed upon by Tyrannosaurus, though it is less certain that two adults did battle in the fanciful manner often depicted in museum displays and popular images. The functions of the frills and three distinctive facial horns on its head have long inspired debate. Traditionally, these have been viewed as defensive weapons against predators. More recent interpretations find it probable that these features were primarily used in species identification, courtship, and dominance display, much like the antlers and horns of modern ungulates.

Triceratops was traditionally placed within the "short-frilled" ceratopsids, but modern cladistic studies show it to be a member of the Chasmosaurinae which usually have long frills. Two species, T. horridus and T. prorsus, are considered valid today, from the seventeen species that have ever been named. Research published in 2010 concluded that the contemporaneous Torosaurus, a ceratopsid long regarded as a separate genus, represents Triceratops in its mature form. This view has been disputed; further data is needed to settle the debate.

Triceratops has been documented by numerous remains collected since the genus was first described in 1889 by American paleontologist Othniel Charles Marsh. Specimens representing life stages from hatchling to adult have been found. As the archetypal ceratopsid, Triceratops is one of the most popular dinosaurs, and has been featured in film, postal stamps, and many other types of media.[1]

The first named fossil specimen now attributed to Triceratops is a pair of brow horns attached to a skull roof, found by George Lyman Cannon near Denver, Colorado, in the spring of 1887.[2] This specimen was sent to Marsh, who believed that the formation from which it came dated from the Pliocene, and that the bones belonged to a particularly large and unusual bison, which he named Bison alticornis.[2][3] He realized that there were horned dinosaurs by the next year, which saw his publication of the genus Ceratops from fragmentary remains,[4] but he still believed B. alticornis to be a Pliocene mammal. It took a third and much more complete skull to change his mind.


Illustration of specimen YPM 1871E, the horn cores that were erroneously attributed to Bison alticornis, the first named specimen of Triceratops
Type specimen YPM 1820 of the type species, T. horridus
1896 skeletal restoration of T. prorsus by O. C. Marsh, based on the holotype skull YPM 1822 and referred elements
T. prorsus, Carnegie Museum of Natural History
The skull (AMNH 5116) of this T. horridus composite specimen was formerly assigned to T. elatus
Size comparison with T. horridus in blue and T. prorsus in red
Skull of Triceratops horridus from the Houston Museum of Natural Science
Life reconstruction of a subadult Triceratops horridus
Closeup of pelvis of Triceratops specimen at the Oxford University Museum of Natural History
Specimen nicknamed "Lane", was the most complete known specimen until 2014
First mounted T. horridus skeleton (the holotype of T. "obtusus"), nicknamed "Hatcher", Smithsonian Museum
Yoshi's Trike, an immature specimen with 115 cm horn cores, on display in the Museum of the Rockies in Montana, USA
Skull of specimen DMNH 48617 from the Laramie Formation of eastern Colorado. Based on the age of the formation, it may be the oldest Triceratops known.
A Triceratops mounted next to a Tyrannosaurus at the Los Angeles Natural History Museum
Close up of the jaws and teeth
Front view of skull with a prominent epoccipital fringe, Houston Museum of Natural Science
Examples of periosteal reactive bone in selected specimens of Triceratops
Juvenile and adult skulls—the juvenile skull is about the size of an adult human head
Skull growth series
A, Triceratops prorsus holotype YPM 1822 and B, Torosaurus latus ANSP 15192
Comparisons between the skulls of Triceratops and Nedoceratops
Pie chart of the time averaged census for large-bodied dinosaurs from the entire Hell Creek Formation in the study area
Restoration of Triceratops and smaller Leptoceratops in the Hell Creek environments
Triceratops and other animals of the Hell Creek Formation
1901 illustration by Charles R. Knight